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After fending off the threat of a Neuroi invasion of Romagna and destroying the enemy's nest over Venezia, Yoshika Miyafuji goes back to her home town in the Empire of Fusou. Despite the loss of her magical and healing abilities, the former officer of the 501st Joint Fighter Wing wants to continue studying medicine. This is in order to help those in need, both civilians and those on the front lines alike. She receives an invitation from a prestigious school in Europe and decides to accept the offer, embarking on a journey back to the war-torn continent. However, a new danger arises, as more and more Neuroi manage to break through human defense lines, appearing unnoticed in supposedly safe areas and disturbing the balance of power in Europe. Yoshiki's sense of duty collides with her inability to pilot a Striker Unit, but with her friends in peril, it seems that there is only one thing she can do.

Preparations for a new offensive against the Neuroi—a mysterious race of alien invaders—are well underway. The objective is securing Berlin, the capital city of the Empire of Karlsland, which is necessary for wiping out the Neuroi threat from Europe. However, as the enemy is capable of adapting to the battlefield on a daily basis, the allied forces and the current state of Striker technology might not be enough to achieve a victory. Meanwhile, Yoshika Miyafuji, a Witch from Fuso, continues her medical studies in Lausanne. Having recovered from a recent incident that deprived her of magical power, she is eager to assist in the war effort. The call to arms soon arrives and the scattered witches of the 501st Joint Fighter Wing must be gathered once again for a final push against the enemy. [Written by MAL Rewrite]

Operation Victory Arrow is set in the period between the end of Operation Mars in the second television series and the 2012 Strike Witches movie. The three 30-minute short stories depict the lives of the members who returned to their respective homes. (Source: ANN)

Yoshika Miyafuji joins the 501st Joint Fighter Wing, a defense unit specializing in taking down alien beings known as the Neuroi. However, in days of no invasion, she and the members of the unit are taking down day to day tasks like cooking and laundry. (Source: MAL News)